Ownership Profile

Shareholder Breakdown

Shareholder Holders Value ($MM) % O/S Shares
Institution 39 22.08 39.06 18,902,048
Mutual Fund 12 3.35 2.57 1,242,374
Insider -- -- -- --

Institutional Shareholder Rotation

Shareholder Holders Value Chg ($MM) % O/S Share Change
Buyers 16 7.93 14.46 6,995,961
Sellers 13 -0.89 1.77 -858,454
Net -- 7.04 12.69 6,137,507

Institutional Shareholder Concentration

Value ($MM) % O/S Shares
Total Inst. 22.08 39.06 18,902,048
Top 10 Inst. 21.30 37.50 18,144,986
Top 20 Inst. 21.95 38.81 18,781,977
Top 50 Inst. 22.08 39.06 18,902,048

Institutional Shareholder Style

Holders Value ($MM) % Inst Shares Shares
Growth 3 2.01 6.26 3,026,876
Core Growth 2 0.03 0.06 26,876
Growth 1 1.98 6.20 3,000,000
Income 1 -- 0.01 3,758
Income Value 1 -- 0.01 3,758
Index 6 1.06 2.08 1,007,201
Other 13 14.62 21.19 10,253,886
Broker Dealer 7 0.10 0.20 96,546
Hedge Fund 5 9.43 10.99 5,315,749
VC/Private Equi 1 5.08 10.01 4,841,591
Value 6 0.01 0.03 12,629
Core Value 3 -- 0.01 3,758
Deep Value 3 0.01 0.02 8,871

Top Holders: Institutional Investor

Fund Shares Held % O/S Share Change Filing Date
Vanguard Total Stock Market Index Fund 691,408 1.43 415,351 4/30/18
Vanguard Extended Market Index Fund 438,753 0.91 282,600 4/30/18
iShares Micro-Cap ETF 36,231 0.07 1,152 4/30/18
Fidelity Extended Market Index Fund 33,461 0.07 -- 3/31/18
Fidelity NASDAQ Composite Index Fund 16,656 0.03 -- 3/31/18
Fidelity Total Market Index Fund 6,105 0.01 -- 3/31/18
PowerShares Wilshire Micro-Cap Portfolio 5,359 0.01 595 3/31/18
iShares Core S&P Total U.S. Stock Market ETF 4,270 0.01 70 4/30/18
Vanguard Balanced Index Fund 3,500 0.01 -- 4/30/18
1290 VT Micro Cap Portfolio 2,745 0.01 -- 3/31/18
Legg Mason Small-Cap Quality Value ETF 2,486 0.01 2,486 8/31/17
Vanguard US Equity Index Fund 1,400 -- -- 3/31/18

Top Holders

Holder Shares Held % O/S Share Change Filing Date
Third Rock Ventures, LLC 4,841,591 10.01 -- 3/31/18
Clairmark Investments, Ltd. 3,582,328 7.40 -- 10/31/17
Empery Asset Management, L.P. 3,000,000 6.20 3,000,000 11/1/17
Kingdon Capital Management, L.L.C. 2,213,980 4.58 1,008,055 4/3/18
Highbridge Capital Management, LLC 1,593,626 3.29 1,593,626 3/31/18
Sabby Management, LLC 1,226,117 2.53 877,224 3/31/18
BlackRock Institutional Trust Company, N.A. 442,288 0.91 -430 3/31/18
The Vanguard Group, Inc. 437,110 0.90 110,227 3/31/18
Celniker (Abbie C) 409,915 0.85 -- 10/31/17
Hurly (Stephen A.) 398,031 0.82 -- 10/31/17
Millennium Management LLC 245,265 0.51 245,265 3/31/18
Lynch (Daniel S) 70,879 0.15 -- 10/31/17
Geode Capital Management, L.L.C. 56,222 0.12 -- 3/31/18
Virtu Americas LLC 45,804 0.09 -381,244 3/31/18
DRW Securities, LLC 40,000 0.08 40,000 3/31/18
Squarepoint Capital LLP 38,277 0.08 38,277 3/31/18
Parallax Volatility Advisers, L.P. 36,761 0.08 -- 3/31/18
BNY Mellon Asset Management North America Corporation 36,601 0.08 -- 3/31/18
Simplex Trading, LLC 34,429 0.07 34,429 3/31/18
Tubridy (Karen L) 32,753 0.07 -- 10/31/17
Northern Trust Investments, Inc. 29,621 0.06 9,028 3/31/18
Fiduciary Trust Company 20,000 0.04 -- 3/31/18
Private Advisor Group LLC 19,524 0.04 19,524 3/31/18
Susquehanna Financial Group, LLLP 14,897 0.03 14,897 3/31/18
Morgan Stanley Wealth Management 6,876 0.01 5,000 3/31/18
McCabe (John J) 5,930 0.01 -- 10/31/17
Invesco PowerShares Capital Management LLC 5,359 0.01 -- 3/31/18
BNY Mellon Asset Management 3,758 0.01 -- 3/31/18
IHT Wealth Management, LLC 3,240 0.01 -- 3/31/18
Royce & Associates, LP 2,886 0.01 400 3/31/18
AXA Rosenberg Investment Management LLC 2,745 0.01 -- 3/31/18
BlackRock Investment Management, LLC 2,745 0.01 -- 3/31/18
Advantage Investment Management, LLC 1,000 -- -- 3/31/18
Citi Investment Research (US) 600 -- -- 3/31/18
RBC Capital Markets Wealth Management 500 -- -- 3/31/18
Morgan Stanley & Co. LLC 315 -- -9,871 3/31/18
Cutler Group, LP 61 -- -- 3/31/18
RBC Capital Partners 13 -- 8 3/31/18
Barclays Capital Inc. 1 -- 1 3/31/18

* Insider values reflect direct beneficial ownership.

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